Tafsir of An-Nahl 16:49

ولله يسجد ما في السماوات وما في اﻻرض من دآبة

And to Allah prostrate all that are in the heavens and all that are in the earth, of the moving creatures, As Allah says:

ولله يسجد من فى السموت واﻻرض طوعا وكرها وظللهم بالغدو واﻻصال

And to Allah (alone) all who are in the heavens and the earth fall in prostration, willingly or unwillingly, and so do their shadows in the mornings and in the afternoons. (13:15)

والمﻼيكة وهم ﻻ يستكبرون

and the angels, and they are not proud. means, they prostrate to Allah and are not too proud to worship Him